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Holotype Specimens at the Raymond M. Alf Museum of Paleontology

The Alf Museum is home to the holotype specimens for 18 different species. The collection’s historical strengths have been in fossil footprints – thus, we currently hold 13 ichnotypes, from birds, camels, bear-dogs, and even a spider. More recent work has emphasized bony elements, with several mammalian holotypes as well as a dinosaur.

The links below lead to information on each type specimen, including photographs, bibliographies, and proposed synonymies. Only holotypes and lectotypes, but not paratypes, are listed here.

Ichnotypes

Classification   Ichnogenus Ichnospecies Ichnotype Specimen
Arachnida   Octopodichnus raymondi RAM 139
Aves   Anatipeda alfi RAM 111
    Anatipeda californica RAM 115
    Avipeda gryponyx RAM 110
Mammalia Amphicyonidae Hirpexipes alfi
RAM 100
 
  Camelidae Dizygopodium dorydium RAM 209
  Camelidae Dizygopodium quadracordatum
RAM 216
 
  Camelidae Lamaichnum alfi RAM 159
  Camelidae Lamaichnum macropodum
RAM 165
 
  Creodonta? Quiritipes impendens
RAM 154
 
  Felidae Felipeda bottjeri
RAM 103
 
  Felidae Pycnodactylopus achras RAM 205
  Ursidae Platykopus ilyacalcator RAM 277

Bony Holotypes

Classification   Genus Species Type Specimen
Dinosauria Hadrosauridae Gryposaurus monumentensis
RAM 6797
 
Mammalia Arctocyonidae Mimotricentes tedfordi
RAM 6908
 
  Cylindrodontidae Cylindrodon nebraskensis
RAM 7009
 
  Equidae Megahippus mckennai
RAM 910
 
  Periptychidae Goleroconus alfi
RAM 7171
 


Goleroconus alfi
RAM 7171

Specimen Description
Incomplete right dentary with p4, m1, m2, and part of the trigonid of m3.

Classification
Mammalia: Eparctocyona: Periptychidae

Locality
RAM locality V200120 (Kern County, California)

Geological Horizon
Goler Formation

Geological Age
Tiffanian NALMA, Late Paleocene

Original Publication
McKenna, M. C., Honey, J. G., and Lofgren, D. L. 2008. Goleroconus alfi, a new small periptychid (Mammalia, Eparctocyona) from the late Paleocene of California. Science Series, Los Angeles County Museum 41:29-42.

Current Taxonomic Status
Valid.
 

 

Megahippus mckennai
RAM 910

Specimen Description
Partial skull.

Classification
Mammalia: Perissodactyla: Equidae

Locality
RAM locality V94294 (San Bernardino County, California)

Geological Horizon
Barstow Formation

Geological Age
Barstovian NALMA, Middle Miocene

Original Publication
Tedford, R. H., and R. M. Alf. 1962. A new Megahippus from the Barstow Formation, San Bernardino County, California. Bulletin of the Southern California Academy of Science 61:113–122.

Current Taxonomic Status
Valid.
 

 

Lamaichnum alfi
RAM 159

Specimen Description
Footprint in relief.

Classification
Mammalia: Artiodactyla: Camelidae

Locality
RAM locality V94276 (San Bernardino County, California)

Geological Horizon
Barstow Formation

Geological Age
Barstovian NALMA, Middle Miocene

Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 1999. Camel and horse footprints from the Miocene of California. San Bernadino County Museum Association Quarterly 46:3-19.

Other Important Publications
Lucas, S. G., and A. P. Hunt. 2007. Ichnotaxonomy of camel footprints. New Mexico Museum of Natural History and Science Bulletin 42:155-168. [designates RAM 159 as lectotype for Lamaichnum alfi; synonymizes the species with Lamaichnum guanicoe]

Current Taxonomic Status
Synonymized with Lamaichnum guanicoe (see Lucas and Hunt, 2007).

Notes
Three specimens were designated as the syntype of Lamaichnum alfi by Sarjeant and Reynolds (1999): RAM 159, 166, and 182. Lucas and Hunt (2007), noting that the ICZN rules require a single holotype specimen, nominated RAM 159 as the lectotype for the ichnospecies.

 

Lamaichnum macropodum
RAM 165

Specimen Description
Footprint from pes.

Classification
Mammalia: Artiodactyla: Camelidae

Locality
RAM locality V94215 (San Bernardino County, California)

Geological Horizon
Tecopa Formation

Geological Age
Clarendonian NALMA, Late Miocene

Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 1999. Camel and horse footprints from the Miocene of California. San Bernadino County Museum Association Quarterly 46:3-19.

Other Important Publications
Lucas, S. G., and A. P. Hunt. 2007. Ichnotaxonomy of camel footprints. New Mexico Museum of Natural History and Science Bulletin 42:155-168. [designates RAM 165 as lectotype for Lamaichnum macropodum]

Current Taxonomic Status
Valid.

Notes
Two specimens were designated as the syntype of Lamaichnum alfi by Sarjeant and Reynolds (1999): RAM 146 and 165. Lucas and Hunt (2007), noting that the ICZN rules require a single holotype specimen, nominated RAM 165 as the lectotype for the ichnospecies.
 

 

Cylindrodon nebraskensis
RAM 7009

Specimen Description
Partial right mandible with p4, m1, and m2 in place.

Classification
Mammalia: Rodentia: Cylindrodontidae

Locality
RAM locality V200130 (Sioux County, Nebraska)

Geological Horizon
Chadron Formation

Geological Age
Chadronian NALMA: Late Eocene

Original Publication
Hough, J., and R. Alf. 1956. A Chadron mammalian fauna from Nebraska. Journal of Paleontology 30:132-140.

Current Taxonomic Status
Valid.

Notes
The specimen number was given as “Webb School Museum No. 108” (which was misprinted as Webb School Museum No. 1908 in the caption for figures 1 and 2, p. 133) by Hough and Alf (1956).
 

 

Mimotricentes tedfordi
RAM 6908

Specimen Description
Partial maxilla with P4-M3.

Classification
Mammalia: Ungulata: Arctocyonidae.

Locality
RAM locality V94133 (Kern County, California)

Geological Horizon
Goler Formation

Geological Age
Tiffanian NALMA, Paleocene

Original Publication
McKenna, M. C., and D. L. Lofgren. 2003. Mimotricentes tedfordi, a new arctocyonid from the Late Paleocene of California. Bulletin of the American Museum of Natural History 279:632-643.

Current Taxonomic Status
Valid.
 

 

Gryposaurus monumentensis
RAM 6797

Specimen Description
Nearly complete skull.

Classification
Dinosauria: Ornithischia: Hadrosauridae.

Locality
RAM locality V200218 (Garfield Co., Utah).

Geological Horizon
Kaiparowits Formation.

Geological Age
Late Campanian, Late Cretaceous.

Original Publication
Gates, T. A., and S. D. Sampson. 2007. A new species of Gryposaurus (Dinosauria: Hadrosauridae) from the late Campanian Kaiparowits Formation, southern Utah, USA. Zoological Journal of the Linnean Society 151:351-376.

Current Taxonomic Status
Valid.
 

 

Octopodichnus raymondi
RAM 139

Specimen Description
Part and counterpart slab with trackway.

Classification
Arthropoda: Arachnida

Locality
RAM locality V94004 (Yavapai Co., Arizona).

Geological Horizon
Coconino Formation.

Geological Age
Early Permian.

Original Publication
Sadler, C.J. 1993. Arthropod trace fossils from the Permian De Chelly Sandstone, northeastern Arizona. Journal of Paleontology 67:240-249.

Current Taxonomic Status
Valid.

Notes
The original specimen number was JF 5905.


 

 

Quiritipes impendens
RAM 154

Specimen Description
Trackway.

Classification
Mammalia: Carnivora: ?Creodonta

Locality
RAM locality V94207 (Carbon Co., Wyoming).

Geological Horizon
probably Wasatch Formation

Geological Age
probably Wasatchian NALMA, Early Eocene

Original Publication
Sarjeant, W. A. S., R. E. Reynolds, and M. M. Kissell-Jones. 2002. Fossil creodont and carnivore footprints from California, Nevada, and Wyoming, in Reynolds, R. E., Between the Basins: Exploring the western Mojave and southern Basin and Range Province: Fullerton, California State University, Desert Studies Consortium, pp. 37-50.

Current Taxonomic Status
Valid.
 

 

Hirpexipes alfi
RAM 100

Specimen Description
Trackway.

Classification
Mammalia: Carnivora: Amphicyonidae

Locality
RAM locality V94272 (San Bernardino Co., California).

Geological Horizon
Barstow Formation.

Geological Age
Barstovian NALMA, Miocene.

Original Publication
Sarjeant, W. A. S., R. E. Reynolds, and M. M. Kissell-Jones. 2002. Fossil creodont and carnivore footprints from California, Nevada, and Wyoming, in Reynolds, R. E., Between the Basins: Exploring the western Mojave and southern Basin and Range Province: Fullerton, California State University, Desert Studies Consortium, pp. 37-50.

Current Taxonomic Status
Valid.
 

 

Felipeda bottjeri
RAM 103

Specimen Description
?Right pes print.

Classification

Mammalia: Carnivora: Felidae

Locality
RAM locality V94276 (San Bernardino Co., California).

Geological Horizon
Barstow Formation.

Geological Age
Barstovian NALMA, Miocene.

Original Publication
Sarjeant, W. A. S., R. E. Reynolds, and M. M. Kissell-Jones. 2002. Fossil creodont and carnivore footprints from California, Nevada, and Wyoming, in Reynolds, R. E., Between the Basins: Exploring the western Mojave and southern Basin and Range Province: Fullerton, California State University, Desert Studies Consortium, pp. 37-50.

Current Taxonomic Status
Valid.
 

 

Avipeda gryponyx
RAM 110

Specimen Description
Trackway with seven footprints.

Classification
Aves

Locality
RAM locality V94021 (San Bernardino Co., California).

Geological Horizon
Avawatz Formation.

Geological Age
Clarendonian NALMA, Late Miocene.

Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 2001. Bird footprints from the Miocene of California, in Reynolds, R. E., ed., The changing face of the east Mojave Desert: Fullerton, California State University, Desert Studies Consortium, p. 21-40.

Current Taxonomic Status
Valid.


 

 

Anatipeda californica
RAM 115

Specimen Description
Left pes print.

Classification
Aves

Locality
RAM locality V94135 (San Bernardino Co., California).

Geological Horizon
Avawatz Formation.

Geological Age
Clarendonian NALMA, Late Miocene.

Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 2001. Bird footprints from the Miocene of California, in Reynolds, R. E., ed., The changing face of the east Mojave Desert: Fullerton, California State University, Desert Studies Consortium, p. 21-40.

Current Taxonomic Status
Valid.


 

 

Anatipeda alfi
RAM 111

Specimen Description
Left and right pedal prints.

Classification
Aves

Locality
RAM locality V94135 (San Bernardino Co., California).

Geological Horizon
Avawatz Formation.

Geological Age
Clarendonian NALMA, Late Miocene.

Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 2001. Bird footprints from the Miocene of California, in Reynolds, R. E., ed., The changing face of the east Mojave Desert: Fullerton, California State University, Desert Studies Consortium, p. 21-40.

Current Taxonomic Status
Valid.


 

 

Dizygopodium dorydium
RAM 209

Specimen Description
manus print

Classification
Mammalia: Artiodactyla: Camelidae

Locality
RAM locality V94021 (San Bernardino Co., California).

Geological Horizon
Avawatz Formation.

Geological Age
Clarendonian NALMA, Late Miocene.

Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 1999. Camel and horse footprints from the Miocene of California. San Bernadino County Museum Association Quarterly 46:3-19.

Other Important Publications
Lucas, S. G., and A. P. Hunt. 2007. Ichnotaxonomy of camel footprints. New Mexico Museum of Natural History and Science Bulletin 42:155-168. [designates RAM 209 as lectotype for Dizygopodium dorydium; synonymizes the species with Lamaichnum guanicoe]

Current Taxonomic Status
Synonymized with Lamaichnum guanicoe (see Lucas and Hunt, 2007).

Notes
Two specimens were designated as the syntype of Dizygopodium dorydium by Sarjeant and Reynolds (1999): RAM 197 and 209. Lucas and Hunt (2007), noting that the ICZN rules require a single holotype specimen, nominated the middle print in a series of three in RAM 197 as the lectotype for the ichnospecies.


 

 

Dizygopodium quadracordatum
RAM 216

Specimen Description
slab with prints from left manus and right pes

Classification
Mammalia: Artiodactyla: Camelidae

Locality
RAM locality V94021 (San Bernardino Co., California).

Geological Horizon
Avawatz Formation.

Geological Age
Clarendonian NALMA, Late Miocene.

Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 1999. Camel and horse footprints from the Miocene of California. San Bernadino County Museum Association Quarterly 46:3-19.

Other Important Publications
Lucas, S. G., and A. P. Hunt. 2007. Ichnotaxonomy of camel footprints. New Mexico Museum of Natural History and Science Bulletin 42:155-168. [designates RAM 159 as lectotype for Lamaichnum alfi; synonymizes the species with Lamaichnum guanicoe]

Current Taxonomic Status
Synonymized with Lamaichnum guanicoe (see Lucas and Hunt, 2007).

Notes
Two specimens were designated as the syntype of Lamaichnum alfi by Sarjeant and Reynolds (1999): RAM 197 and 209. Lucas and Hunt (2007), noting that the ICZN rules require a single holotype specimen, nominated RAM 159 as the lectotype for the ichnospecies.
 

 

Pycnodactylopus achras
RAM 205

Specimen Description
Slab with right manus and left pes? prints.

Classification
Mammalia: Carnivora: Felidae

Locality
RAM locality V94134 (San Bernardino Co., California).

Geological Horizon
Avawatz Formation.

Geological Age
Clarendonian NALMA, Late Miocene.

Original Publication
Sarjeant, W. A. S., R. E. Reynolds, and M. M. Kissell-Jones. 2002. Fossil creodont and carnivore footprints from California, Nevada, and Wyoming, in Reynolds, R. E., Between the Basins: Exploring the western Mojave and southern Basin and Range Province: Fullerton, California State University, Desert Studies Consortium, pp. 37-50.

Current Taxonomic Status
Valid.


 

 

Platykopus ilyacalcator
RAM 277

Specimen Description
Manus print.

Classification
Mammalia: Carnivora: Ursidae

Locality
RAM locality V94163 (Clark Co., Nevada).

Geological Horizon
Muddy Creek Formation.

Geological Age
Hemphillian NALMA, Late Miocene.

Original Publication
Sarjeant, W. A. S., R. E. Reynolds, and M. M. Kissell-Jones. 2002. Fossil creodont and carnivore footprints from California, Nevada, and Wyoming, in Reynolds, R. E., Between the Basins: Exploring the western Mojave and southern Basin and Range Province: Fullerton, California State University, Desert Studies Consortium, pp. 37-50.

Current Taxonomic Status
Valid.


 

 

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