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Digging for fossils in the Barstow Formation
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Dr. Don Lofgren
Dr. Andrew Farke
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Goler Formation
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Holotype Specimens at the Raymond M. Alf Museum of
Paleontology
The Alf Museum is home to the holotype specimens for 18 different species.
The collection’s historical strengths have been in fossil footprints – thus,
we currently hold 13 ichnotypes, from birds, camels, bear-dogs, and even a
spider. More recent work has emphasized bony elements, with several
mammalian holotypes as well as a dinosaur.
The links below lead to information on each type specimen, including
photographs, bibliographies, and proposed synonymies. Only holotypes and
lectotypes, but not paratypes, are listed here.Ichnotypes
| Classification |
|
Ichnogenus |
Ichnospecies |
Ichnotype Specimen |
| Arachnida |
|
Octopodichnus |
raymondi |
RAM 139
 |
| Aves |
|
Anatipeda |
alfi |
RAM 111
 |
| |
|
Anatipeda |
californica |
RAM 115
 |
| |
|
Avipeda |
gryponyx |
RAM 110
 |
| Mammalia |
Amphicyonidae |
Hirpexipes |
alfi |
RAM 100
|
| |
Camelidae |
Dizygopodium |
dorydium |
RAM 209
 |
| |
Camelidae |
Dizygopodium |
quadracordatum |
RAM 216
|
| |
Camelidae |
Lamaichnum |
alfi |
RAM 159
 |
| |
Camelidae |
Lamaichnum |
macropodum |
RAM 165
|
| |
Creodonta? |
Quiritipes |
impendens |
RAM 154
|
| |
Felidae |
Felipeda |
bottjeri |
RAM 103
|
| |
Felidae |
Pycnodactylopus |
achras |
RAM 205
 |
| |
Ursidae |
Platykopus |
ilyacalcator |
RAM 277
 |
Bony Holotypes
| Classification |
|
Genus |
Species |
Type Specimen |
| Dinosauria |
Hadrosauridae |
Gryposaurus |
monumentensis |
RAM 6797
|
| Mammalia |
Arctocyonidae |
Mimotricentes |
tedfordi |
RAM 6908
|
| |
Cylindrodontidae |
Cylindrodon |
nebraskensis |
RAM 7009
|
| |
Equidae |
Megahippus |
mckennai |
RAM 910
|
| |
Periptychidae |
Goleroconus |
alfi |
RAM 7171
|
Goleroconus alfi
RAM 7171
Specimen Description
Incomplete right dentary with p4, m1, m2, and part of the trigonid of m3.
Classification
Mammalia: Eparctocyona: Periptychidae
Locality
RAM locality V200120 (Kern County, California)
Geological Horizon
Goler Formation
Geological Age
Tiffanian NALMA, Late Paleocene
Original Publication
McKenna, M. C., Honey, J. G., and Lofgren, D. L. 2008. Goleroconus alfi,
a new small periptychid (Mammalia, Eparctocyona) from the late Paleocene of
California. Science Series, Los Angeles County Museum 41:29-42.
Current Taxonomic Status
Valid.
Megahippus
mckennai
RAM 910
Specimen Description
Partial skull.
Classification
Mammalia: Perissodactyla: Equidae
Locality
RAM locality V94294 (San Bernardino County, California)
Geological Horizon
Barstow Formation
Geological Age
Barstovian NALMA, Middle Miocene
Original Publication
Tedford, R. H., and R. M. Alf. 1962. A new Megahippus from the
Barstow Formation, San Bernardino County, California. Bulletin of the
Southern California Academy of Science 61:113–122.
Current Taxonomic Status
Valid.
Lamaichnum alfi
RAM 159
Specimen Description
Footprint in relief.
Classification
Mammalia: Artiodactyla: Camelidae
Locality
RAM locality V94276 (San Bernardino County, California)
Geological Horizon
Barstow Formation
Geological Age
Barstovian NALMA, Middle Miocene
Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 1999. Camel and horse footprints
from the Miocene of California. San Bernadino County Museum Association
Quarterly 46:3-19.
Other Important Publications
Lucas, S. G., and A. P. Hunt. 2007. Ichnotaxonomy of camel footprints. New
Mexico Museum of Natural History and Science Bulletin 42:155-168.
[designates RAM 159 as lectotype for Lamaichnum alfi; synonymizes the
species with Lamaichnum guanicoe]
Current Taxonomic Status
Synonymized with Lamaichnum guanicoe (see Lucas and Hunt, 2007).
Notes
Three specimens were designated as the syntype of Lamaichnum alfi by
Sarjeant and Reynolds (1999): RAM 159, 166, and 182. Lucas and Hunt (2007),
noting that the ICZN rules require a single holotype specimen, nominated RAM
159 as the lectotype for the ichnospecies.

Lamaichnum
macropodum
RAM 165
Specimen Description
Footprint from pes.
Classification
Mammalia: Artiodactyla: Camelidae
Locality
RAM locality V94215 (San Bernardino County, California)
Geological Horizon
Tecopa Formation
Geological Age
Clarendonian NALMA, Late Miocene
Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 1999. Camel and horse footprints
from the Miocene of California. San Bernadino County Museum Association
Quarterly 46:3-19.
Other Important Publications
Lucas, S. G., and A. P. Hunt. 2007. Ichnotaxonomy of camel footprints. New
Mexico Museum of Natural History and Science Bulletin 42:155-168.
[designates RAM 165 as lectotype for Lamaichnum macropodum]
Current Taxonomic Status
Valid.
Notes
Two specimens were designated as the syntype of Lamaichnum alfi by
Sarjeant and Reynolds (1999): RAM 146 and 165. Lucas and Hunt (2007), noting
that the ICZN rules require a single holotype specimen, nominated RAM 165 as
the lectotype for the ichnospecies.
Cylindrodon
nebraskensis
RAM 7009
Specimen Description
Partial right mandible with p4, m1, and m2 in place.
Classification
Mammalia: Rodentia: Cylindrodontidae
Locality
RAM locality V200130 (Sioux County, Nebraska)
Geological Horizon
Chadron Formation
Geological Age
Chadronian NALMA: Late Eocene
Original Publication
Hough, J., and R. Alf. 1956. A Chadron mammalian fauna from Nebraska.
Journal of Paleontology 30:132-140.
Current Taxonomic Status
Valid.
Notes
The specimen number was given as “Webb School Museum No. 108” (which was
misprinted as Webb School Museum No. 1908 in the caption for figures 1 and
2, p. 133) by Hough and Alf (1956).
Mimotricentes
tedfordi
RAM 6908
Specimen Description
Partial maxilla with P4-M3.
Classification
Mammalia: Ungulata: Arctocyonidae.
Locality
RAM locality V94133 (Kern County, California)
Geological Horizon
Goler Formation
Geological Age
Tiffanian NALMA, Paleocene
Original Publication
McKenna, M. C., and D. L. Lofgren. 2003. Mimotricentes tedfordi, a
new arctocyonid from the Late Paleocene of California. Bulletin of the
American Museum of Natural History 279:632-643.
Current Taxonomic Status
Valid.
Gryposaurus
monumentensis
RAM 6797
Specimen Description
Nearly complete skull.
Classification
Dinosauria: Ornithischia: Hadrosauridae.
Locality
RAM locality V200218 (Garfield Co., Utah).
Geological Horizon
Kaiparowits Formation.
Geological Age
Late Campanian, Late Cretaceous.
Original Publication
Gates, T. A., and S. D. Sampson. 2007. A new species of Gryposaurus (Dinosauria:
Hadrosauridae) from the late Campanian Kaiparowits Formation, southern Utah,
USA. Zoological Journal of the Linnean Society 151:351-376.
Current Taxonomic Status
Valid.
Octopodichnus
raymondi
RAM 139
Specimen Description
Part and counterpart slab with trackway.
Classification
Arthropoda: Arachnida
Locality
RAM locality V94004 (Yavapai Co., Arizona).
Geological Horizon
Coconino Formation.
Geological Age
Early Permian.
Original Publication
Sadler, C.J. 1993. Arthropod trace fossils from the Permian De Chelly
Sandstone, northeastern Arizona. Journal of Paleontology 67:240-249.
Current Taxonomic Status
Valid.
Notes
The original specimen number was JF 5905.

Quiritipes
impendens
RAM 154
Specimen Description
Trackway.
Classification
Mammalia: Carnivora: ?Creodonta
Locality
RAM locality V94207 (Carbon Co., Wyoming).
Geological Horizon
probably Wasatch Formation
Geological Age
probably Wasatchian NALMA, Early Eocene
Original Publication
Sarjeant, W. A. S., R. E. Reynolds, and M. M. Kissell-Jones. 2002. Fossil
creodont and carnivore footprints from California, Nevada, and Wyoming, in
Reynolds, R. E., Between the Basins: Exploring the western Mojave and
southern Basin and Range Province: Fullerton, California State University,
Desert Studies Consortium, pp. 37-50.
Current Taxonomic Status
Valid.
Hirpexipes alfi
RAM 100
Specimen Description
Trackway.
Classification
Mammalia: Carnivora: Amphicyonidae
Locality
RAM locality V94272 (San Bernardino Co., California).
Geological Horizon
Barstow Formation.
Geological Age
Barstovian NALMA, Miocene.
Original Publication
Sarjeant, W. A. S., R. E. Reynolds, and M. M. Kissell-Jones. 2002. Fossil
creodont and carnivore footprints from California, Nevada, and Wyoming, in
Reynolds, R. E., Between the Basins: Exploring the western Mojave and
southern Basin and Range Province: Fullerton, California State University,
Desert Studies Consortium, pp. 37-50.
Current Taxonomic Status
Valid.
Felipeda bottjeri
RAM 103
Specimen Description
?Right pes print.
Classification
Mammalia: Carnivora: Felidae
Locality
RAM locality V94276 (San Bernardino Co., California).
Geological Horizon
Barstow Formation.
Geological Age
Barstovian NALMA, Miocene.
Original Publication
Sarjeant, W. A. S., R. E. Reynolds, and M. M. Kissell-Jones. 2002. Fossil
creodont and carnivore footprints from California, Nevada, and Wyoming, in
Reynolds, R. E., Between the Basins: Exploring the western Mojave and
southern Basin and Range Province: Fullerton, California State University,
Desert Studies Consortium, pp. 37-50.
Current Taxonomic Status
Valid.
Avipeda gryponyx
RAM 110
Specimen Description
Trackway with seven footprints.
Classification
Aves
Locality
RAM locality V94021 (San Bernardino Co., California).
Geological Horizon
Avawatz Formation.
Geological Age
Clarendonian NALMA, Late Miocene.
Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 2001. Bird footprints from the
Miocene of California, in Reynolds, R. E., ed., The changing face of the
east Mojave Desert: Fullerton, California State University, Desert Studies
Consortium, p. 21-40.
Current Taxonomic Status
Valid.

Anatipeda
californica
RAM 115
Specimen Description
Left pes print.
Classification
Aves
Locality
RAM locality V94135 (San Bernardino Co., California).
Geological Horizon
Avawatz Formation.
Geological Age
Clarendonian NALMA, Late Miocene.
Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 2001. Bird footprints from the
Miocene of California, in Reynolds, R. E., ed., The changing face of the
east Mojave Desert: Fullerton, California State University, Desert Studies
Consortium, p. 21-40.
Current Taxonomic Status
Valid.

Anatipeda alfi
RAM 111
Specimen Description
Left and right pedal prints.
Classification
Aves
Locality
RAM locality V94135 (San Bernardino Co., California).
Geological Horizon
Avawatz Formation.
Geological Age
Clarendonian NALMA, Late Miocene.
Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 2001. Bird footprints from the
Miocene of California, in Reynolds, R. E., ed., The changing face of the
east Mojave Desert: Fullerton, California State University, Desert Studies
Consortium, p. 21-40.
Current Taxonomic Status
Valid.

Dizygopodium
dorydium
RAM 209
Specimen Description
manus print
Classification
Mammalia: Artiodactyla: Camelidae
Locality
RAM locality V94021 (San Bernardino Co., California).
Geological Horizon
Avawatz Formation.
Geological Age
Clarendonian NALMA, Late Miocene.
Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 1999. Camel and horse footprints
from the Miocene of California. San Bernadino County Museum Association
Quarterly 46:3-19.
Other Important Publications
Lucas, S. G., and A. P. Hunt. 2007. Ichnotaxonomy of camel footprints. New
Mexico Museum of Natural History and Science Bulletin 42:155-168.
[designates RAM 209 as lectotype for Dizygopodium dorydium;
synonymizes the species with Lamaichnum guanicoe]
Current Taxonomic Status
Synonymized with Lamaichnum guanicoe (see Lucas and Hunt,
2007).
Notes
Two specimens were designated as the syntype of Dizygopodium
dorydium by Sarjeant and Reynolds (1999): RAM 197 and 209. Lucas and
Hunt (2007), noting that the ICZN rules require a single holotype specimen,
nominated the middle print in a series of three in RAM 197 as the lectotype
for the ichnospecies.

Dizygopodium quadracordatum
RAM 216
Specimen Description
slab with prints from left manus and right pes
Classification
Mammalia: Artiodactyla: Camelidae
Locality
RAM locality V94021 (San Bernardino Co., California).
Geological Horizon
Avawatz Formation.
Geological Age
Clarendonian NALMA, Late Miocene.
Original Publication
Sarjeant, W. A. S., and R. E. Reynolds. 1999. Camel and horse footprints
from the Miocene of California. San Bernadino County Museum Association
Quarterly 46:3-19.
Other Important Publications
Lucas, S. G., and A. P. Hunt. 2007. Ichnotaxonomy of camel footprints. New
Mexico Museum of Natural History and Science Bulletin 42:155-168.
[designates RAM 159 as lectotype for Lamaichnum alfi; synonymizes the
species with Lamaichnum guanicoe]
Current Taxonomic Status
Synonymized with Lamaichnum guanicoe (see Lucas and Hunt, 2007).
Notes
Two specimens were designated as the syntype of Lamaichnum alfi
by Sarjeant and Reynolds (1999): RAM 197 and 209. Lucas and Hunt (2007),
noting that the ICZN rules require a single holotype specimen, nominated RAM
159 as the lectotype for the ichnospecies.
Pycnodactylopus
achras
RAM 205
Specimen Description
Slab with right manus and left pes? prints.
Classification
Mammalia: Carnivora: Felidae
Locality
RAM locality V94134 (San Bernardino Co., California).
Geological Horizon
Avawatz Formation.
Geological Age
Clarendonian NALMA, Late Miocene.
Original Publication
Sarjeant, W. A. S., R. E. Reynolds, and M. M. Kissell-Jones. 2002. Fossil
creodont and carnivore footprints from California, Nevada, and Wyoming, in
Reynolds, R. E., Between the Basins: Exploring the western Mojave and
southern Basin and Range Province: Fullerton, California State University,
Desert Studies Consortium, pp. 37-50.
Current Taxonomic Status
Valid.

Platykopus
ilyacalcator
RAM 277
Specimen Description
Manus print.
Classification
Mammalia: Carnivora: Ursidae
Locality
RAM locality V94163 (Clark Co., Nevada).
Geological Horizon
Muddy Creek Formation.
Geological Age
Hemphillian NALMA, Late Miocene.
Original Publication
Sarjeant, W. A. S., R. E. Reynolds, and M. M. Kissell-Jones. 2002. Fossil
creodont and carnivore footprints from California, Nevada, and Wyoming, in
Reynolds, R. E., Between the Basins: Exploring the western Mojave and
southern Basin and Range Province: Fullerton, California State University,
Desert Studies Consortium, pp. 37-50.
Current Taxonomic Status
Valid.


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